Drainage divide networks – Part 2: Response to perturbations

Drainage divides are organized into tree-like networks that may record information about drainage divide mobility. However, views diverge about how to best assess divide mobility. Here, we apply a new approach of automatically extracting and ordering drainage divide networks from digital elevation models to results from landscape evolution model experiments. We compared landscapes perturbed by strike-slip faulting and spatiotemporal variations in erodibility to a reference model to assess which topographic metrics (hillslope relief, flow distance, and χ ) are diagnostic of divide mobility. Results show that divide segments that are a minimum distance of ∼ 5 km from river confluences strive to attain constant values of hillslope relief and flow distance to the nearest stream. Disruptions of such patterns can be related to mobile divides that are lower than stable divides, closer to streams, and often asymmetric in shape. In general, we observe that drainage divides high up in the network, i.e., at great distances from river confluences, are more susceptible to disruptions than divides closer to these confluences and are thus more likely to record disturbance for a longer time period. We found that across-divide differences in hillslope relief proved more useful for assessing divide migration than other tested metrics. However, even stable drainage divide networks exhibit across-divide differences in any of the studied topographic metrics. Finally, we propose a new metric to quantify the connectivity of divide junctions.


Introduction
Divide migration is a time-dependent process that is difficult to quantify. While the effects of regional-scale drainage captures may be preserved within sedimentary archives (e.g., Clift et al., 2006), this is unlikely for smaller-scale drainage captures or gradual divide migration. In such cases, most studies rely on topographic indicators. Mobile divides are typically inferred from post-drainage-capture evidence: distorted drainage structures, low divides (wind gaps), or high tributary junction angles (e.g., Clark et al., 2004) (Fig. 1). However, divide mobility may also be expressed in the topography without major drainage captures or flow reversals but as a result of the more gradual migration of divides. Willett et al. (2014) inferred drainage divide mobility from across-divide differences in χ values, a proxy for steadystate river channel elevation (Perron and Royden, 2013).
They argued that changes in drainage area within mountain ranges, e.g., due to tectonic strain of the crust (Yang et al., 2015), may commonly lead to relative differences in incision rate and the formation of low-relief landscapes that are bordered by migrating divides. Whipple et al. (2017a, b), however, argued that the timescale of such changes is too short to profoundly affect mountainous landscapes. Instead, they argued that transient low-relief landscapes, such as those in southeastern Tibet, are more likely to be formed by regional changes in rock uplift rate and upstream propagation of knickpoints between the adjusted and unadjusted parts of landscapes. They also cast doubt on the ease of comparing across-divide differences in drainage network geometry (i.e., χ values) where the common base level is far and opposing rivers may incise into areas of different rock types, different rock uplift rates, or different climates. Whipple et al. (2017a,  b) instead proposed that the shape of drainage divides themselves holds clues about their mobility (Fig. 1). Amongst the topographic parameters that they tested are across-divide differences in channel elevation at a reference drainage area, mean headwater hillslope gradient, and mean headwater local relief.
In summary, several different metrics have been proposed that may allow for the quantification of divide mobility in both natural and modeled landscapes. Forte and Whipple (2018) compared the performance of these metrics with a landscape evolution model in which they induced divide mobility and concluded that across-divide differences in relief or gradient better depict divide motion than χ . In their analysis, however, they focused on divide motion that is perpendicular to the regional drainage direction and averaged divide migration rates as well as topographic metrics across the entire width of the model domain so that each time step is associated with single values for divide migration rate, erosion rate difference, and the tested topographic metrics. In part 1 of this study (Scherler and Schwanghart, 2020a), we presented a new approach for the automatic identification and ordering of drainage divide networks in a digital elevation model (DEM), which removes the necessity of manually selecting drainage divides for comparison. Here, we present experiments with a numerical landscape evolution model that we conducted to investigate how drainage divide networks respond to different perturbations, including fault activity and differences in erodibility. In contrast to previous studies that examined the response of drainage divides to perturbations, we studied the entire drainage divide network in an objective manner and examined how different portions of the divide network respond to perturbations. In addition, we tested the utility of a new metric that quantifies the connectivity of drainage divide junctions.

Landscape evolution model
We studied the response of divide networks to stream captures and divide migration using the TopoToolbox Landscape Evolution Model (TTLEM; Campforts et al., 2017). In our experiments, we modeled the topographic evolution of a 20 km × 20 km square block (50 m node spacing) subject to uniform rock uplift, stream-power-based fluvial incision (e.g., Howard and Kerby, 1983;Whipple and Tucker, 1999), and hillslope diffusion (e.g., Culling, 1963): where z is elevation (L), t is time (T), U is rock uplift rate (L T −1 ), A is upstream area (L 2 ), S is local channel slope (L L −1 ), K r is a parameter of the efficiency of river incision (T −1 ) (K r = 1 × 10 −5 yr −1 ), and m and n are dimensionless constants with values of 0.5 and 1, respectively. The last term on the right-hand side depicts elevation change due to the divergence in diffusive hillslope transport q s (L 3 L −1 T −1 ), which we consider to be a linear function of hillslope gradient: q s = −D∇z, where D is the diffusivity (L 2 T −1 ) of soil creep (D = 2 × 10 −3 m 2 yr −1 ). All four edges of the block were fixed in elevation (z = 0 m), which forced rivers to flow outwards. The uplift rate (U = 1 mm yr −1 ) was constant in all models. Our choice of parameter values was guided by the study of Whipple et al. (2017b), who tested a wide range of rock uplift and erosional efficiency parameters and found almost no difference of divide mobility in models with and without hillslope diffusion and for n values of 1 and 2. We started from a flat surface with imposed random noise and ran the experiment for 30 Myr until the topography reached a steady state. The result of this model, which we termed "Initialize", provided the starting point for four other models that we ran for 10 Myr (Fig. 2). The model "Reference" included no further changes. In the model "Rotating", we included a circular (10 km diameter) left-lateral strikeslip fault that was active throughout the experiment. Strikeslip faults are well known for enforcing drainage captures and thus divide mobility (e.g., Castelltort et al., 2012;Duval and Tucker, 2015). Although the rotating block has, to our knowledge, no real-world equivalent, this model setup represents a convenient way of simulating extended periods of strike-slip faulting, as the fault does not intersect the model boundary (Braun and Sambridge, 1997). The fault slip rate was fixed at 4 mm yr −1 , which corresponds to an angular velocity of 8 × 10 −7 rad yr −1 , resulting in ∼ 460 • of total rotation during the model run. We note that the rotating movement requires interpolation and thus leads to numerical diffusion of elevations within the rotating disk. However, the resulting change in total volume by interpolation is < 0.03 % of the volume uplifted during the same time and therefore small. The model "Inclined" included 1 km thick and 5 km spaced layers of 50 % reduced erosional efficiency of rivers (K r ), dipping 30 • towards northwest. The Inclined model is representative of a landscape in which rivers incise into tilted sedimentary rocks of nonuniform rock strength, similar to what has been studied by Forte and Whipple (2018). During the experiment, the combination of surface lowering and inclination resulted in the resistant layers regularly sweeping from southeast to northwest across the simulated landscape. The model "Spheres" included 30 randomly assembled spheres of 3 km diameter with 75 % reduced erosional efficiency of rivers (K r ). This experiment may represent incision of a region that is characterized by country rocks with more resistant magmatic intrusions. The expected behavior of this model is similar to the landscape response to localized perturbations studied by O'Hara et al. (2018).

Topographic analysis
We analyzed the modeled topography and the associated drainage divide network. For each modeled topography and at each time step (dt = 40 000 years), we first computed flow directions and flow accumulation, and we subsequently identified the stream network using a drainage area threshold of 0.2 km 2 . We next derived the drainage divide network on the basis of the stream network and using the algorithm proposed in Scherler and Schwanghart (2020a). We calculated divide distances and divide orders based on the Topo ordering scheme (Scherler and Schwanghart, 2020a). As topographic metrics, we included elevation (z), hillslope relief (HR), and horizontal flow distance to the stream network (FD). HR was measured as the elevation difference between a point on the divide and the nearest river location as measured by the distance along local flow directions. We also computed χ values on either side of a divide using a reference area A 0 of 1 m 2 , a reference concavity θ ref of 0.45, and setting the base level x b to 0 at the edge of the model domain (e.g., Perron and Royden, 2013): For each divide edge, we computed these topographic metrics and the erosion rate (Eq. 1) for the two neighboring pixels that belong to adjacent drainage basins and denoted the across-divide minimum, maximum, sum, difference, and average in any one metric X as X min , X max , X, X, and X, respectively. Erosion rates were based on the erosion rate of the first downslope stream pixel to reduce the impact of local noise along hillslopes. Topographic metrics of entire divide segments are based on those of the divide edges that it is composed of. For quantifying across-divide differences in topographic metrics and erosion rates, irrespective of the actual values, we used normalized indices of the form X/ X. One such index that we frequently used in our study is the divide asymmetry index (DAI = | HR/ HR|), which is the absolute value of the normalized hillslope relief difference and which ranges between 0 (symmetric) and 1 (most asymmetric).
The above-described across-divide differences in topographic metrics essentially aim to quantify divide mobility. In contrast, Spotila (2012) studied the stability of divides and argued that divide junctions and pyramidal peaks are more stable than solitary linear divides and might therefore act as anchor points for drainage divide networks. He proposed that divide junctions are more difficult to erode than linear divides due to their greater volume of topography per unit area, their greater mechanical stability, and their reduction of confluent flows (Spotila, 2012). He also suggested that the stability of divide junctions is related to the number of joining drainage divides. Because the divide junctions obtained from our algorithm cannot connect more than four divide segments (Scherler and Schwanghart, 2020a) -and most often connect three segments -we introduce a new metric to quantify divide junction connectivity, C J : We define C J to correspond to the sum of the ratios of the Euclidean distance, d, and the divide distance, d d , of all divide edges, n, within a specified maximum divide distance, d d,max , times the ratio of the cell size, dx, and d d,max . The dimensionless quantity C J is sensitive to the number of divides within a given divide distance from a junction weighted by their orientation towards the junction (Fig. 3). The value d d,max reflects the divide distance over which differences in junction connectivity are measured. For junctions that connect a constant number of straight and infinitely long divide segments, C J is not sensitive to the value of d d,max . However, for actual junctions, C J is typically sensitive to the value of d d,max because as d d,max grows, increasingly more junctions are at a distance d d < d d,max of a specific junction, and thus the number of divide segments grows with d d (Fig. 3a). In general, C J will be sensitive to the position of a junction within the drainage divide network if the junction's maximum divide distance from an endpoint is smaller than d d,max . In other cases, C J will provide a measure of how connected a junction is within a network or, in other words, how prominent the junction is compared to other junctions in the network. In this study, we used a d d,max value of 5 km.

General behavior
The simulated landscapes, along with their drainage divide networks at the end of the numerical experiments, are shown in Figs. 4 and 5, and in the "Video supplement" (Scherler and Schwanghart, 2020b) we provide movies of all simulations.
To provide a measure of the mobility of drainage divides, we computed the percentages of drainage area that were exchanged during the simulations between individual catchments that drain to the margin of the model domain (Fig. 6).
Except for the Reference model, all models are characterized by notable changes in drainage area and mobile drainage divides. Area changes in the Initialize model are large in the beginning but level off rapidly during the first 1 Myr. Although area changes are small after 1 Myr, they continue for another 20 Myr, during which they are mostly decreasing. In the Rotating model, large area changes appear as discrete pulses induced by drainage captures of major streams (Fig. 5b), whereas the background area changes during rotation and faulting are relatively small (< 0.1 % per 40 kyr). Area changes in the Inclined model are moderate (∼ 0.25 % per 40 kyr) throughout the simulation and oscillate in conjunction with the passage of more resistant layers through the landscape (Fig. 5c). Area changes in the Spheres model are generally more pronounced if the resistant spheres appear in the course of rivers, which forces them to steepen and to induce surface uplift upstream as opposed to their appearance at drainage divides, which increases the height of the divide but does not induce drainage divide migration at a larger scale (Fig. 5d).

Network topology
We first analyzed the response of the entire drainage network topology to the perturbations by quantifying the aggregated length of divide segments as a function of their order (Fig. 7). The first few million years of the Initialize model are characterized by large changes in divide lengths and orders. Initially, the divide network extends to orders as high as 100 but rapidly contracts as the drainage network becomes dendritic. After about 5 Myr, the highest orders are down to 60. Subsequent changes result in some scatter of the divide lengths but not in the range of divide orders. Compared to the Reference model, in which the divide network structure no longer changes, the Rotating, Inclined, and Spheres models exhibit changes in the divide network, mostly at divide orders greater than ∼ 20. This observation is related to the fact that loworder divides are distributed across the entire model domain and their number is accordingly high. Any of the perturbations we imposed only affect some of these divides, and thus the impact on their average length is rather small. In contrast, high-order divides are constrained to the highest parts of the modeled land surface and their numbers are much lower. The imposed perturbations typically affect a greater portion of them and hence the scatter in divide lengths is wider. In the Rotating and Spheres models, we also observed that maximum divide orders occasionally extend to higher values, but these changes are rather small. We note that the above observations also prevail when considering divide distance instead of divide order because the two are linearly related (Scherler and Schwanghart, 2020a), with divide distance ∼ 430 m × divide order. The 430 m corresponds to the mean length of the divide segments.

Topographic metrics
We next studied how the above-described disturbances affect drainage divide metrics during the simulations (Fig. 8).
For all models, we computed the averages of the topographic parameters measured at drainage divides of specific divide distance intervals (Fig. 8a-d). As in the analysis of dividesegment lengths by order, it should be kept in mind that the numbers of divide segments, or their aggregated lengths (Fig. 7), are much higher for low orders and distances compared to higher ones. For reference, a divide order of 20 corresponds to a divide distance of approximately 9 km. In the Initialize model, all of the studied metrics attain a constant value that remains unchanged in the Reference model (Fig. 8) and that may or may not depend on the divide distance. For example, the mean elevation and junction connectivity (C J ) clearly increase with divide distance, whereas the flow distance exhibits only minor dependence on divide distance, and hillslope relief appears unrelated to divide distance. The dependency of some metrics on divide distance is partly explained by the model setup. Although divides with low distances also occur at higher elevation, the bulk of them are near the model edge, close to zero elevation. In contrast, divides at high distances are exclusively found near the center of the model, where elevations are also high. Similarly, the junction connectivity (C J ) is high in the model center, where the divides are far from most of the endpoints, which are more abundant near the edges of the model. It is also worth noting that none of the normalized acrossdivide differences in the topographic metrics attain zero values in the Reference model. This means that even at topographic steady state, there are residual across-divide differences in hillslope relief, flow distance, and χ. In the case of χ, these also depend on the divide distance and are greater closer to the model edge, where divide distances are low. In the perturbed models, we observed fluctuations in all topographic metrics, although of different magnitudes. For example, a comparison of across-divide differences in erosion rate with differences in hillslope relief, flow distance, and χ ( Fig. 8e-h) shows that the normalized difference in hillslope relief (i.e., the divide asymmetry index) is sensitive to drainage divide mobility in all perturbed models, whereas across-divide differences in χ and flow distance are sensitive to divide mobility in the Rotating model but less so in the Inclined and Spheres models. The junction connectivity (C J ) metric attains temporally averaged values in the perturbed models that are quite similar to the constant values in the Reference model. In many cases, the deviations from the Reference model are greater the higher up in the divide network, i.e., for higher divide distances. This pattern is particularly visible in the Inclined model, wherein the amplitudes of the oscillations in all of the parameters increase with divide distance.

Minimum hillslope relief and flow distance
Motivated by the observation of constant values in hillslope relief and flow distance in the Reference model, as well as in actual landscapes (Scherler and Schwanghart, 2020a), and by our expectation that small values in either one would be found where one catchment loses area to another (Fig. 1), we next compared how minimum hillslope relief (HR min ), minimum flow distance (FD min ), and the divide asymmetry index (DAI) vary with divide distance in the Reference model and the three models with landscape perturbances (Fig. 9). In contrast to the average values in Fig. 8, we provide these metrics for all divide edges during the last 1 Myr of the model runs. Note also that we plotted data points in an order that brings high DAI values to the front to better assess where asymmetric divides are located, but in all four models, relatively high DAI points may plot on top of low DAI points. In the Reference model, both minimum hillslope relief and minimum flow distance reach relatively steady values (HR min ∼ 250-350 m; FD min ∼ 400-600 m) at a divide distance of ∼ 5 km. At lower divide distances, both HR min and FD min approach zero -simply because divides are defined to start at the stream network -and these divides can become increasingly more asymmetric. It is notable, however, that some of the highest HR min and FD min values are also observed at low divide distances of approximately 1-2 km. In the three other models, the transition between quite variable divides at short distances and more steady "background" values at higher distances appears to be preserved, but we observe generally more variability. For example, in the Rotating model, we observe divides with significantly lower HR min and FD min values at higher distances. These divides are particularly prominent at a distance of ∼ 10 and ∼ 15-22 km and correspond to the position of the strike-slip fault. Where HR min and FD min are low, DAI values are relatively high (divides are highly asymmetric), although there are also divides that have high DAI but regular HR min and FD min values. In the Inclined and Spheres models, the HR min and FD min values are never as low as in the Rotating model at divide distances > 5 km, which reflects the lack of drainage captures. Instead, we observe frequent excursions to both higher and lower HR min and FD min values, either across all divide distances (Inclined) or at specific locations (Spheres). Deviations from the average values in the Reference model are greatest in the Spheres model compared to all other perturbed models and always correspond to peaks that grow where strong spheres are exhumed. In the three disturbance models, DAI values are generally higher than in the Reference model -although divides with high HR min and FD min values and at great divide distances mostly appear to have somewhat lower DAI values.

Junction connectivity
The spatial pattern of divide junction connectivity (C J ) values at the end of the landscape evolution experiments     Fig. 10) partly follows the pattern of divide distances (Fig. 4). Junctions with higher C J values tend to occur at higher elevation and at greater divide distance (d d ). In the Reference model, the highest C J values occupy the center of the model domain and the centers of the four quadrants of the model domain, resembling the five on six-sided dice. In the Rotating model, these clusters of high C J values are maintained, but their connection is disrupted by the strikeslip fault, which induces low C J values. The centrally located divide junctions occupy a similar range in C J values but are all shifted to higher elevations (Fig. 4). Similar, although lower, offsets to higher elevations occur in the Inclined and Spheres models, where junctions coincide with rocks of reduced erodibility. In the Spheres model, the basic structure of C J values is similar to that of the Reference model, but the highest C J values are steered towards the less erodible spheres, whereby they also attain C J values that are distinctly higher than in any of the other models (Fig. 10b). In general, divide junctions with combinations of elevation and C J values that are outside the range of values observed in the Reference model are found in the most disturbed parts of the landscape (Fig. 10c). In summary, the perturbed models appear to induce mostly changes in junction elevation, whereas changes in junction connectivity (C J ) are seemingly constrained to the Spheres model.

Quantifying drainage divide mobility
The analysis of stream networks has become a standard tool for inferring tectonic forcing and landscape history (e.g., Wobus et al., 2006;Kirby and Whipple, 2012;Demoulin, 2012;Schwanghart and Scherler, 2014). The divide network holds the potential to record similar tectonic forcing, but also other aspects of landscape history (e.g., Willett et al., 2014). The question is which divide metrics are useful to analyze, and what do they tell us about landscape history? Our Rotating model induced relatively sudden drainage captures (Fig. 6). Because such events are associated with the dissection of drainage divides, reliable indicators are values of hillslope relief (HR) and flow distance (FD) that are much lower compared to the values that divides (>∼ 5 km divide distance) strive for (Fig. 9). More gradual divide migration, however, likely lacks such simple diagnostic criteria, and in those cases, across-divide differences in topographic metrics may be more suitable indicators of divide mobility. The most commonly used metric to infer drainage divide mobility is the across-divide difference in χ (Willett et al., 2014). Although the utility of this metric has recently received some critique (Whipple et al., 2017b;Forte and Whipple, 2018), it has become a popular tool for studying drainage divides. Whipple et al. (2017b) and Forte and Whipple (2018) instead advocated the use of other topographic metrics, including mean gradient, mean local relief, and channel bed eleva-tion, measured at or upstream of a reference drainage area. We note that these latter metrics are typically highly correlated and very similar to the hillslope relief and DAI metrics that we included in this study. Figure 11 shows how normalized across-divide differences in χ, hillslope relief (HR), and flow distance (FD) compare to normalized across-divide differences in erosion rate (ER), evaluated for each divide edge from the last 1 Myr of the landscape evolution experiments Reference, Rotating, Inclined, and Spheres. We find that across-divide differences in hillslope relief (HR; Fig. 11a) are most sensitive to the disturbances included in the models, whereas across-divide differences in χ are similarly sensitive to disturbances in the Rotating model but less so in the Inclined and Spheres models (Figs. 11b, 8). Across-divide differences in flow distance (FD) are the least sensitive to disturbances in the models and show the largest scatter when compared with erosion rates (Fig. 11c). However, there is also substantial scatter in the relationship between across-divide differences in hillslope relief and erosion rate, which partly depends on the divide distance. In general, we observe that the scatter is higher for divide distances <∼ 5 km (dark blue in Fig. 11), which corresponds to the value below which we observe large variability in divide morphology, even in the Reference model (Fig. 9). To quantify the correlation of the normalized across-divide differences in topographic metrics with normalized acrossdivide differences in erosion rate, we fitted a linear model to all drainage divide edges from the entire model runs, categorized into 1 km divide distance bins, and show the resulting coefficients of determination (R 2 ) in Fig. 12. As already suspected from Figs. 8 and 11, the R 2 values differ between models and metrics and also depend on divide distance. In general, we observe that all metrics perform poorly at divide distances <∼ 5 km and that across-divide differences in flow distance perform poorly even at higher distances. The highest R 2 values are linked to across-divide differences in hillslope relief, whereas across-divide differences in χ attain similar R 2 values in the Rotating model but some of the lowest R 2 values of all metrics in the Inclined and Spheres model. This difference may be explained by the fact that in the latter two models, we introduced spatial variability in the erosional efficiency of rivers (K r ) that we did not account for in our acrossdivide comparison of χ, as would be required (Willett et al., 2014). In natural landscapes, however, these values and their variability are rarely well known.
We speculate that the influence of divide distance on topographic metric-erosion rate relationships may also account for the differences in scatter observed by Sassolas-Serrayet et al. (2019) in landscape evolution experiments similar to our Initialize model between larger and smaller basin areas. But even when excluding divides of low order or low divide distance, we still observe considerable scatter in the topographic metric-erosion rate relationships, which, at the very least, demands caution when interpreting divide morphology in terms of mobility. In this regard, studying Fig. 5 Figure 11. Relationship between across-divide differences in the topographic metrics hillslope relief (HR), chi (χ), and flow distance (FD), with across-divide differences in erosion rates (ERs) for all divide edges in the last 1 Myr of the landscape evolution experiments Reference, Rotating, Inclined, and Spheres. Marker color denotes the divide distance (d d ). Note that the markers are sorted by their divide distance to show the highest divide distance above data points with lower divide distance, and data points with lower divide distance occur beneath those with higher divide distance. Figure 12. Coefficient of determination for linear regressions between normalized across-divide differences in different topographic metrics (HR: hillslope relief; FD: flow distance; χ : chi) and normalized across-divide differences in erosion rates as a function of drainage divide distance in the three perturbation models. D. Scherler and W. Schwanghart: Drainage divide networks -Part 2 of the landscape evolution experiments (see the "Video supplement"; Scherler and Schwanghart, 2020b) is insightful: where drainage divides are migrating, one typically observes a range of across-divide topographic metric values that vary considerably during the migration. In other words, despite a continuous divide migration at a large scale, there is often small-scale variability in divide morphology that may in part be related to across-divide differences in topographic metrics lagging behind across-divide differences in erosion rate.

and the videos
As a final note, we emphasize that the above observations are from our numerical experiments, which depict an idealized world. It is clear that the complexities present in nature, such as anisotropic and variable rock properties, hydroclimatic gradients, mass-wasting events, and biological influences on erosion processes and rates, can lead to landscape patterns that bias any of the above topographic metrics and need to be taken into account when inferring divide dynamics from divide metrics in natural landscapes.

Divide network dynamics
Stream networks tend to attain configurations that are in equilibrium with the geological and climatic environment, given an initial condition (e.g., Rinaldo et al., 2014). Because drainage divides are defined by adjacent drainage basins, the geometry of divide networks should attain a similar equilibrium, which expresses itself in both the geometry of divides and the topology of divide networks. Our numerical experiments have shown that during the initial establishment of a stream network, on a relatively flat surface, both stream and divide networks are far from their steady-state configuration and characterized by networks that extend to high orders (Fig. 7) and long divide distances. During the subsequent extension and shrinkage of individual streams towards their steady-state configuration, the divide network contracts and primarily high-order divide segments shorten and become fewer, whereas divides of low orders maintain their frequencies (Fig. 7).
In general, divide segments of high order, i.e., at great distance from endpoints, appear to be the most responsive to landscape disturbances (Fig. 8). In the case of the Rotating model, this is in part expected because the inner rotating part of the landscape contains the highest-order divide segments (Fig. 4b). In the cases of the Inclined and Spheres models, it may be related to the increased probability of recording a disturbance because the adjoining basins cover a larger area compared to lower-order divides. In other words, if drainage captures happen somewhere within a drainage basin, this will most likely influence divides further upstream. Over a distance of less than ∼ 5 km from divide network endpoints, the divide segments transition from low interfluves at river junctions to high topographic ridges, as seen in the Reference model (Fig. 9). In the other models, most of the investigated morphometric parameters are quite variable over the same distance and can be seen to rapidly adjust to disturbances such as drainage captures or migrating divides. Such behavior is consistent with the observation that the timescale of a river's response to changes in drainage area increases with the distance from the divide to the outlet of a river (Whipple et al., 2017b). To reliably distinguish the morphologic effects of real disturbances from "noise" close to the river, a minimum divide distance of perhaps ∼ 5 km, as in our analysis of the Big Tujunga divide network (Scherler and Schwanghart, 2020a), appears appropriate. This minimum divide distance could be lower or higher, depending on factors like drainage density and average hillslope relief, for example.
Our new junction connectivity index (C J ) complements existing topographic metrics in assessing divide network dynamics. For example, the junction connectivity in our Rotating model is low along the fault (Fig. 10), consistent with the absence of stable divides. In the Spheres model, however, the appearance of more resistant rocks at the surface often resulted in the migration of divides towards the spheres (Fig. 5c, "Video supplement"; Scherler and Schwanghart, 2020b). In this case, parts of the drainage divide network were mobile, not stable, but they moved towards particularly stable portions in the landscape. Therefore, the junction connectivity index (C J ) may also be interpreted as an attractor or centrality index (Phillips et al., 2015) that quantifies how strong a drainage divide network has been pulled towards and anchored at a certain junction (Spotila, 2012).

Conclusions
Based on landscape evolution model experiments in which we forced divides to migrate, we found that stable drainage divides strive to attain a constant hillslope relief and flow distance from the nearest stream, provided a sufficiently large divide distance to avoid confounding influences near the edges of the divide network. In our experiments this distance is ∼ 5 km from endpoints. Simple indicators of mobile divides are anomalously low hillslope relief or flow distance values, which could signal beheaded valleys or future capture events. Overall, drainage divides located high up in the network, i.e., at great distance from endpoints, are more vulnerable than divides closer to endpoints of the network and are more likely to record disturbance for a longer time period. In our comparison of different topographic metrics to assess drainage divide mobility, we found that across-divide differences in hillslope relief proved more useful for assessing divide migration than other tested metrics.
Code availability. The divide algorithm developed in Scherler and Schwanghart (2020a) has been implemented in the TopoToolbox v2 (Schwanghart and Scherler, 2014). The codes will be made available with the next TopoToolbox release and shall be accessible at https://github.com/wschwanghart/topotoolbox (last access: 17 April 2020).