Bartholdy, J., Ernstsen, V. B., Flemming, B. W., Winter, C., Bartholomä,
A., and Kroon, A.: On the formation of current ripples, Sci. Rep., 5, 11390, https://doi.org/10.1038/srep11390, 2015.
Blanchard, G. F., Guarini, J.-M., Gros, P., and Richard, P.: Seasonal effect
on the relationship between the photosynthetic capacity of intertidal
microphytobenthos and temperature, J. Phycol., 33, 723–728,
https://doi.org/10.1111/j.0022-3646.1997.00723.x, 1997.
Borchers, H. W.: pracma: Practical Numerical Math Functions, available at:
https://cran.r-project.org/web/packages/pracma/index.html (last access: 8 April 2021), 2019.
Brakenhoff, L., Schrijvershof, R., van der Werf, J., Grasmeijer, B., Ruessink, G., and van der Vegt, M.: From Ripples to Large-Scale Sand
Transport: The Effects of Bedform-Related Roughness on Hydrodynamics and
Sediment Transport Patterns in Delft3D, J. Mar. Sci. Eng., 8, 1–25, https://doi.org/10.3390/jmse8110892, 2020.
Cheng, C. H.: NIOZ racetrack sand-shell experiment, data belonging to the paper: Sediment shell-content diminishes current-driven sand ripple development and migration, 4TU.ResearchData [data set], https://doi.org/10.4121/12852113, 2021.
Cheng, C. H., Soetaert, K., and Borsje, B. W.: Sediment Characteristics over
Asymmetrical Tidal Sand Waves in the Dutch North Sea, J. Mar. Sci. Eng., 8, 1–16, https://doi.org/10.3390/jmse8060409, 2020.
Curran, J. C.: An investigation of bed armoring process and the formation of
microclusters, in: 2nd Joint Federal Interagency Conference, Las Vegas, 1–12, 2010.
Damveld, J. H., Reijden, K. J., Cheng, C., Koop, L., Haaksma, L. R., Walsh,
C. A. J., Soetaert, K., Borsje, B. W., Govers, L. L., Roos, P. C., Olff, H.,
and Hulscher, S. J. M. H.: Video transects reveal that tidal sand waves affect the spatial distribution of benthic organisms and sand ripples,
Geophys. Res. Lett.,45, 11837–11846, https://doi.org/10.1029/2018GL079858, 2018.
Damveld, J. H., Roos, P. C., Borsje, B. W., and Hulscher, S. J. M. H.:
Modelling the two-way coupling of tidal sand waves and benthic organisms: A
linear stability approach, Environ. Fluid Mech., 19, 1073–1103, https://doi.org/10.1007/s10652-019-09673-1, 2019.
Dey, S.: Incipient Motion of Bivalve Shells on Sand Beds under Flowing Water, J. Eng. Mech., 129, 232–240, https://doi.org/10.1061/(ASCE)0733-9399(2003)129:2(232), 2003.
Dietrich, W., Kirchner, J., Ikeda, H., and Iseya, F.: Sediment Supply and
Development of Coarse Surface Layer in Gravel Bedded Rivers, Nature, 340,
215–217, https://doi.org/10.1038/340215a0, 1989.
Earle, S.: Sea-Floor Sediments, available at:
https://geo.libretexts.org/@go/page/7876 (last access: 8 April 2021), 2020.
Friedrichs, M., Graf, G., and Springer, B.: Skimming flow induced over a
simulated polychaete tube lawn at low population densities, Mar. Ecol. Prog.
Ser., 192, 219–228, https://doi.org/10.3354/meps192219, 2000.
Friedrichs, M., Leipe, T., Peine, F., and Graf, G.: Impact of macrozoobenthic
structures on near-bed sediment fluxes, J. Mar. Syst., 75, 336–347,
https://doi.org/10.1016/j.jmarsys.2006.12.006, 2009.
Friend, P. L., Lucas, C. H., Holligan, P. M., and Collins, M. B.: Microalgal
mediation of ripple mobility, Geobiology, 6, 70–82,
https://doi.org/10.1111/j.1472-4669.2007.00108.x, 2008.
Gornitz, V.: Encyclopedia of Paleoclimatology and Ancient Environments, Springer, https://doi.org/10.1007/978-1-4020-4411-3, 2008.
Gutiérrez, J., Jones, C., Strayer, D., and Iribarne, O.: Mollusks as
ecosystem engineers: The role of shell production in aquatic habitats, Oikos, 101, 79–90, https://doi.org/10.1034/j.1600-0706.2003.12322.x, 2003.
Herman, P., Middelburg, J., and Heip, C.: Benthic community structure and
sediment processes on an intertidal flat: Results from the ECOFLAT project,
Cont. Shelf Res., 21, 2055–2071, https://doi.org/10.1016/S0278-4343(01)00042-5, 2001.
Huettel, M. and Rusch, A.: Transport and degradation of phytoplankton in
permeable sediment, Limnol. Oceanogr., 45, 534–549, https://doi.org/10.4319/lo.2000.45.3.0534, 2000.
Idier, D., Astruc, D., and Hulscher, S. J. M. H.: Influence of bed roughness
on dune and megaripple generation, Geophys. Res. Lett., 31, L13214, https://doi.org/10.1029/2004GL019969, 2004.
Kidwell, S. M.: Palaeobiological and sedimentological implications of fossil
concentrations, Nature, 318, 457–460, https://doi.org/10.1038/318457a0, 1985.
Kösters, F. and Winter, C.: Exploring German Bight coastal morphodynamics based on modelled bed shear stress, Geo-Mar. Lett., 34, 21–36, https://doi.org/10.1007/s00367-013-0346-y, 2014.
Langlois, V. and Valance, A.: Initiation and evolution of current ripples on
a flat sand bed under turbulent water flow, Eur. Phys. J. E, 22, 201–208, https://doi.org/10.1140/epje/e2007-00023-0, 2007.
Lapôtre, M., Lamb, M., and McElroy, B.: What sets the size of current
ripples?, Geology, 45, G38598.1, https://doi.org/10.1130/G38598.1, 2017.
Lichtman, I. D., Baas, J. H., Amoudry, L. O., Thorne, P. D., Malarkey, J.,
Hope, J. A., Peakall, J., Paterson, D. M., Bass, S. J., Cooke, R. D., Manning, A. J., Davies, A. G., Parsons, D. R., and Ye, L.: Bedform migration in a mixed sand and cohesive clay intertidal environment and implications for bed material transport predictions, Geomorphology, 315, 17–32,
https://doi.org/10.1016/j.geomorph.2018.04.016, 2018.
Ligges, U., Short, T., Kienzle, P., Schnackenberg, S., Billinghurst, D.,
Borchers, H.-W., Carezia, A., Dupuis, P., Eaton, J. W., Farhi, E., Habel, K., Hornik, K., Krey, S., Lash, B., Leisch, F., Mersmann, O., Neis, P., Ruohio, J., Smith, J. O., Stewart, D., and Weingessel, A.: signal: Signal Processing, available at:
https://cran.r-project.org/web/packages/signal/index.html (last access: 8 April 2021), 2015.
Malarkey, J., Baas, J. H., Hope, J. A., Aspden, R. J., Parsons, D. R., Peakall, J., Paterson, D. M., Schindler, R. J., Ye, L., Lichtman, I. D., Bass, S. J., Davies, A. G., Manning, A. J., and Thorne, P. D.: The pervasive
role of biological cohesion in bedform development, Nat. Commun., 6, 6257,
https://doi.org/10.1038/ncomms7257, 2015.
Miedema, S. and Ramsdell, R.: Hydraulic transport of sand/shell mixtures in
relation with the critical velocity, Terra Aqua, 122, 18–27, 2011.
Mietta, F., Chassagne, C., Manning, A. J., and Winterwerp, J. C.: Influence
of shear rate, organic matter content, pH and salinity on mud flocculation,
Ocean Dynam., 59, 751–763, https://doi.org/10.1007/s10236-009-0231-4, 2009.
Nelson, T. R., Voulgaris, G., and Traykovski, P.: Predicting wave-induced
ripple equilibrium geometry, J. Geophys. Res.-Oceans, 118, 3202–3220,
https://doi.org/10.1002/jgrc.20241, 2013.
Pilditch, C. A., Emerson, C. W., and Grant, J.: Effect of scallop shells and
sediment grain size on phytoplankton flux to the bed, Cont. Shelf Res., 17, 1869–1885, https://doi.org/10.1016/S0278-4343(97)00050-2, 1997.
Pope, N., Widdows, J., and Brinsley, M.: Estimation of bed shear stress using
the turbulent kinetic energy approach – A comparison of annular flume and
field data, Cont. Shelf Res., 26, 959–970, https://doi.org/10.1016/j.csr.2006.02.010,
2006.
Precht, E. and Huettel, M.: Advective pore-water exchange driven by surface
gravity waves and its ecological implications, Limnol. Oceanogr., 48, 1674–1684, https://doi.org/10.4319/lo.2003.48.4.1674, 2003.
Ramsdell, R. and Miedema, S.: Hydraulic transport of sand/shell mixtures, in:
WODCON XIX, WODA – World Organization of Dredging Associations, Beijing, 1–21, 2010.
R Core Team: R: A language and environment for statistical computing, R Foundation for Statistical Computing, Vienna, Austria, available at:
https://www.R-project.org (last access: 8 April 2021), 2020.
Russell-Hunter, W. D. (Ed.): Chapter 1 Overview: Planetary Distribution of and Ecological Constraints upon the Mollusca, in: The Mollusca: Ecology, vol. 6, Academic Press, Orlando, USA, 1–27, 1983.
Seibold, E. and Berger, W. (Eds.): Sources and Composition of Marine Sediments BT – The Sea Floor: An Introduction to Marine Geology, Springer International Publishing, Cham, 45–61, 2017.
Shen, H. W. and Lu, J.: Development and Prediction of Bed Armoring, J. Hydraul. Eng., 109, 611–629, https://doi.org/10.1061/(ASCE)0733-9429(1983)109:4(611), 1983.
Soulsby, R.: Chapter 5 The Bottom Boundary Layer of Shelf Seas, in: Physical
Oceanography of Coastal and Shelf Seas, vol. 35, edited by: Johns, B., Elsevier, New York, USA, 189–266, 1983.
Soulsby, R.: Dynamics of Marine Sands: A manual for Practical Applications,
Thomas Telford Publishing, London, UK, 1997.
Sugiyama, J. and Kobayashi, K.: wvtool: Image Tools for Automated Wood
Identification, available at:
https://cran.r-project.org/web/packages/wvtool/index.html (last access: 8 April 2021), 2016.
Tuijnder, A. P., Ribberink, J. A. N. S., and Hulscher, S. J. M. H.: An
experimental study into the geometry of supply-limited dunes, Sedimentology,
56, 1713–1727, https://doi.org/10.1111/j.1365-3091.2009.01054.x, 2009.
van Ledden, M., van Kesteren, W. G. M., and Winterwerp, J. C.: A conceptual
framework for the erosion behaviour of sand–mud mixtures, Cont. Shelf Res.,
24, 1–11, https://doi.org/10.1016/j.csr.2003.09.002, 2004.
Van Oyen, T., de Swart, H. E., and Blondeaux, P.: Bottom topography and
roughness variations as triggering mechanisms to the formation of sorted
bedforms, Geophys. Res. Lett., 37, 1–5, https://doi.org/10.1029/2010GL043793, 2010.
van Rijn, L. C.: Principles of sediment transport in rivers, estuaries and coastal seas, Aqua Publications Amsterdam, Amsterdam, the Netherlands, 1993.
van Rijn, L. C., Nieuwjaar, M. W. C., van der Kaay, T., Nap, E., and van Kampen, A.: Transport of Fine Sands by Currents and Waves, J. Waterw. Port Coast. Ocean Eng., 119, 123–143, https://doi.org/10.1061/(ASCE)0733-950X(1993)119:2(123), 1993.
Vericat, D., Batalla, R. J., and Garcia, C.: Breakup and reestablishment of
the armour layer in a large gravel-bed river below dams: The lower Ebro,
Geomorphology, 76, 122–136, https://doi.org/10.1016/j.geomorph.2005.10.005, 2006.
Wilcock, P. and Detemple, B.: Persistence of Armor Layers in Gravel-Bed Streams, Geophys. Res. Lett, 32, L08402, https://doi.org/10.1029/2004GL021772, 2005.
Witbaard, R., Bergman, M. J. N., van Weerlee, E., and Duineveld, G. C. A.: An
estimation of the effects of Ensis directus on the transport and burial of
silt in the near-shore Dutch coastal zone of the North Sea, J. Sea Res., 127, 95–104, https://doi.org/10.1016/j.seares.2016.12.001, 2016.
